Collecting duct principal cell (V2 receptor)

Target gland or tissue · V2 cAMP Aquaporin-2

Vasopressin binds the basolateral V2 receptor, raises cAMP and PKA, and shuttles preformed aquaporin-2 vesicles into the apical membrane. It also raises inner medullary urea transport, sharpening the gradient the water follows.

Traced from the start

  1. Supraoptic and paraventricular magnocellular neurones
  2. Posterior pituitary — vasopressin release
  3. Collecting duct principal cell (V2 receptor)

Detail

Channels
AQP2 is apical and regulated; AQP3 and AQP4 are basolateral and constitutive
Range
Urine osmolality can be varied from about 50 to 1200 mOsm/kg
Requires
An intact medullary concentration gradient — loop diuretics abolish it and therefore blunt concentrating ability

When it goes wrong

Long-term lithium

Vasopressin resistance (nephrogenic diabetes insipidus): dilute polyuria that does not respond to desmopressin; amiloride is the usual treatment

Hypercalcaemia or hypokalaemia

Acquired concentrating defect with polyuria, reversible once the electrolyte is corrected

Tolvaptan or demeclocycline

V2 blockade used to raise sodium in SIADH resistant to fluid restriction

Practise this structure

1 question in the bank tagged Water Balance.